Suleiman Abu Ghaith, who married Bin Laden's daughter Fatima, on Wednesday recounted a dramatic meeting with the jubilant al-Qaida chief in an Afghanistan cave complex on the night of September 11, 2001.
"Did you learn what happened? We are the ones who did it," Bin Laden declared, according to Abu Ghaith.
The 48-year-old from Kuwait told the court he warned Bin Laden that he would feel the full force of America's wrath following the attacks on New York and Washington.
Bin Laden replied simply by telling him "You're being too pessimistic."
Within months, the US-led invasion had ousted the Taliban from power in Afghanistan, and Bin Laden was forced onto the run.
A 10-year manhunt ended when the al-Qaida leader was shot dead by US Navy SEALs during a daring raid on his hideout in Pakistan in 2011.
Abu Ghaith had not been expected to testify during his trial where he is charged with conspiracy to kill Americans and conspiracy to support terrorists.
He faces life imprisonment if convicted by a jury at the trial, which is expected to conclude within days.
Speaking in Arabic, translated into English by an interpreter, Abu Ghaith also denied trying to recruit people for al-Qaida, as prosecutors have alleged.
"There is no one recruiting, but Osama Bin Laden. My intention was not recruiting anyone," he said.
And, asked by his lawyer if he ever wanted to kill Americans, he responded "No."
"My intention was to deliver a message I believed in," he said, denouncing the oppression of Muslims.
Presenting himself as an imam, he said he went to Taliban-controlled Afghanistan in June 2001 because he had a "serious desire to get to know the new Islamic government."
His other aim was "teaching and preaching," he said, adding that was something he didn't accomplish.
Clad in a suit with an open-collared shirt, the balding suspect sporting a salt-and-pepper beard admitted having recorded several videos at the request of Bin Laden, who, he said, summoned him after learning he was a Kuwaiti imam.
He said he had never met Richard Reid, a British man who tried to explode a bomb hidden in his shoes on a Paris-Miami flight in December 2001, three months after the 9/11 attacks.
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